Five new species of water mite genus Atractides Koch, 1837 from Bhutan with new records (Acari: Hydrachnidia: Hygrobatidae)
  Torrenticolid water mites of Bhutan. Genera Torrenticola Piersig, 1896 and Neoatractides Lundblad, 1941 (Acari: Hydrachnidia: Torrenticolidae)
  The water mite genus Sperchon Kramer, 1877 in Bhutan (Acari: Hydrachnidia: Sperchontidae), with the description of three new species