A new genus and species of the oribatid mite family, Machuellidae, is described under the name Gredosella fraternalis n. gen., n. sp. The specimen studied was extracted from a soil sample collected in a burned pine forest at Arenas de San Pedro (Sierra de Gredos, Province of Avila, Central Spain). Its most important distinguishing character is that it bears a fused tibia and tarsus on all its legs. Machuella capitata Kuliev, 1967 is transferred to Gredosella.